Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Pennsylvania People

Federal Staff

Duane R. Diefenbach

Duane Diefenbach Unit Leader Website: Tel: (814) 865 - 3992

Upon completion of my Ph.D. I worked for the Pennsylvania Game Commission as their biometrician for 7 years. There I began my current research involving grassland sparrows and black bears. Since 1999 I have been a research scientist with the Pennsylvania Unit and enjoy the challenges in integrating research to improve management decisions.


Tyler Wagner Assistant Unit Leader Tel: (814) 865 - 6592

Office photo

David Walter Assistant Unit Leader Website: Tel: (814) 867 - 4763

After finishing my Ph.D. I served as a post-doctoral Researcher at the National Wildlife Research Center for nearly 5 years. A majority of my research explores spatial ecology of various species throughout the U.S. using GIS to further our understanding of wildlife presence/absence, nutrition, and disease.

University Staff

Alyssia Church

BJ Scovern Tel: (814) 865 - 4511

Students and Post Docs

Danielle Begley-Miller

Danielle Begley-Miller Tel: (814) 865 - 2180

I graduated with my MS in June of 2013 from Wright State University in Dayton, OH. My previous research focused on the impacts of white-tailed deer herbivory on forest understory plant communities in Northern Wisconsin. I'm now in the co-op unit working on The-Deer Forest Study with Duane Diefenbach in pursuit of my PhD in Wildlife. I'm excited about the DFS and its potential to answer much needed questions about the influences of other factors (in combination with deer browsing) on forest vegetation.

Emily Carrollo

Kit fox work in Utah

Chaz Crawford

I received my B.S. in Wildlife Science from Virginia Tech in 2008 where I worked on several non-invasive carnivore projects including using remotely-triggered camera traps to estimate abundance and density of carnivores in eastern rainforests of Madagascar. Since graduation I continued to pursue carnivore research and over the past five years have worked on seven carnivore projects in five states and two countries employing both non-invasive and capture techniques studying species as diverse as kit fox, black bear, fossa, coyote, civets, fishers, and jaguars. Currently I am pursuing my master’s degree with advisor Dr. David Walter focusing on spatial analysis of large mammals to assess harvest vulnerability. When not engaged in my studies you can find me exploring the outdoors and if I’m lucky discovering hidden climbing areas around central PA.

Tyler S. Evans

Tyler Evans Tel: (814) 865 - 4183

Originally from Salem, OH, I earned my B.S. in Wildlife and Fisheries Resources from West Virginia University in May 2012, and I am currently using GIS to develop a surveillance plan for chronic wasting disease (CWD) under the direction of Dr. W. David Walter in the Pennsylvania Cooperative Fish and Wildlife Research Unit. When I am not in the office, I prefer to spend the majority of my time outdoors either hunting in the Ohio hardwoods or fishing. Additionally, I tend to watch my fair share of Mountaineer (WVU) and Nittany Lion football.

Evan Faulk

Evan Faulk

Originally from Poulsbo, WA, I earned a B.S. in Fish & Wildlife Management from Montana State University in May 2012. While attending MSU, I spent my summers working and volunteering in fisheries research throughout the Rocky Mountains, from northern Montana to New Mexico, primarily with native cutthroat trout populations. My coursework and job experience led me to a graduate research position at Penn State with Dr. Tyler Wagner. In close collaboration with the National Park Service, I am studying headwater tributary fish communities within the Eastern Rivers & Mountains Network. When I'm not out chasing fish for research purposes, I'm chasing fish with my fly rod, mountain biking the endless rock gardens of central PA, or playing the guitar.

Nick S. Forman

Nicholas Forman

Before starting in Fall 2012 on my M.S at Penn State, I received my B.S. from Guilford College in Greensboro, North Carolina, where I did an honor’s thesis on the microhabitat affinities of salamanders in impacted landscapes. Field research has taken me to Nicaragua, New Zealand, and Australia, the latter where I spent a year conducting research on a potential biological control for the Cane Toad and contributing to studies on Lumholtz’s Tree Kangaroos. My interests are in spatial ecology and population ecology, with a recent focus on mammals. The project I will be working on with the PA CoOp unit will determine River Otter abundances in Pennsylvania using the non-invasive technique of fecal DNA analysis. Outside of academics, I play rugby, meddle in the culinary arts, and find any excuse to be outside.

Laura Gigliotti

Laura Gigliotti

Originally from Plymouth, MA I graduated from Colby College in Waterville, ME in 2012, where I majored in biology and minored in anthropology. As an undergraduate my research focused on termite diversity and ladybeetle foraging behaviors. I am currently working with Dr. Duane Diefenbach and the PA Cooperative Fish and Wildlife Research Unit on a master’s project looking at snowshoe hare habitat use and mortality rates in response to climate change and forest management in the state. Outside of school I enjoy spending as much time as possible outdoors.

Jason M. Hill

Jason Hill Tel: (814) 865 - 9219

Originally from Marshalltown, Iowa Jason earned his B.S. in Wildlife Biology from the University of Montana in 2001. After a yearlong stint working with sea otters for the USGS in Central California, Jason returned to ornithological field research. He participated in research studies in Costa Rica (house wrens), New York (barn swallows), Florida (red-cockaded woodpeckers), Minnesota (American woodcock), California (cavity-nesting birds), and Maui (po'ouli capture team). He completed his M.S. in Ecology at University of Connecticut under Chris Elphick studying post-fledging ecology of saltmarsh sparrows. He completed his Ph.D. at Penn State in Ecology under Duane Diefenbach studying the population dynamics of grassland sparrows on reclaimed surface mines.

Bill Kanapaux

Bill Kanapaux Tel: (814) 863 - 7544

Megan Kepler

Megan Kepler

My interest in the field of fisheries biology began at a very young age. As I began thinking about my career goals, I wanted to explore the broader field of biology to really get a feeling of how encompassing biology truly was. I attended Lock Haven University and pursued a biology degree with concentrations in cellular and environmental biology. During that time, I was able to complete internships that strengthened my fisheries knowledge, one with the Clinton County Conservation District and a second with the Pennsylvania Fish and Boat Commission. After this point, I knew I wanted to continue my education and also gain more work experience. I began working as a research technician for Dr. Tyler Wagner of the Pennsylvania Cooperative Fish and Wildlife Research Unit where I was able to assist in a wide variety of research projects. Currently I am a pursuing a master’s degree with Dr. Tyler Wagner as my advisor. My thesis work will focus on Comparative Lake Trout Bioenergetics. Additionally, I enjoy spending most of my spare time outside. Whether it is hunting, fishing or just taking a walk with my dogs, my love for the outdoors has always been a large part of my life and has helped develop my career path.

Clayton Lutz

Clayton Lutz

Growing up in central Pennsylvania, I spent most of my childhood exploring the ridges and valleys of the Appalachians. I was fascinated with wildlife and tried to learn anything I could about the plants and animals that I encountered. Consequently, it was a natural fit for me to pursue a career in wildlife. I began by attending Juniata College, where I received a bachelor’s degree in Ecology and Environmental Science. While at Juniata, I conducted research on the nesting ecology of map turtles, and my passion for scientific research and wildlife conservation was formed. After graduation, I worked as a nongame biologist for 4 years with the Arizona Game and Fish Department, where I conducted research on the desert tortoise, Sonoran tiger salamander, and Kanab ambersnail. In 2006, I returned to PA to work as a private consultant performing research on bat migration and habitat use, as well as to administer presence/absence surveys in relation to wind energy development. In 2008, I began working for the PA Game Commission trapping, tagging, and radio-tracking white-tailed deer. I began my graduate studies at Penn State in the spring of 2010. My thesis research topic is female dispersal patterns in white-tailed deer. I am also currently a Wildlife Diversity Biologist with the PA Game Commission. I still get the same thrill out of being outdoors and learning about wildlife but now I enjoy sharing those experiences with my wife, one-year-old daughter, and our energetic chocolate lab.

Steve Midway

Steve Midway Tel: (814) 863 - 2506

I received a BS in Wildlife and Fisheries Biology at the University of Vermont (2005) where I worked as a Fisheries Technician for the Vermont Coop Unit. In 2008, I finished an MS in Fisheries Science at North Carolina State University, again in collaboration with the North Carolina Coop Unit. I spent part of 2009 working for the Florida Fish and Wildlife Conservation Commission before starting a PhD in Marine Biology at the University of North Carolina Wilmington, which was completed in 2013.

William Miller Tel: (717) 327 - 9672

Glenn Stauffer Tel: (841) 865 - 9219

Wendy Vreeland

Wendy Vreeland

As a late bloomer in the field of hunting and fishing, I have been enjoying the outdoors through many different activities beyond my family camping trips as a youngster. Having grown up along the south shore of Long Island, New York, my outdoor experiences included weekend sailing trips to the beach. While in high school, attending an ornithology course changed my life’s path. After completing my Bachelor’s of Science degree in Wildlife Management at the University of Maine, my work experiences have taken me through Maine, Vermont, California and currently Pennsylvania. Aside from hunting and fishing, I spend a lot of time with my husband gardening, making wine and beer, photography, and caring for our English bulldog.

Shannon White

Shannon White

Having earned a B.S. in Biology and Environmental Studies from Randolph-Macon College and a M.S. in Fisheries and Wildlife Science from Virginia Tech, I’ve devoted my academic career to understanding various aspects of fish ecology. My research has included topics on non-game fish conservation, stream habitat restoration, and social learning in salmonids. Through this, I developed a keen interest in determining how the behavior of individual fish drive observed patterns in populations and across landscapes; a topic I will explore further while pursuing a Ph.D. in the Pennsylvania Cooperative Fish and Wildlife Research Unit.


Current Staff

Federal Staff: 3

Masters Students: 8

Phd Students: 4

Post Docs: 4

University Staff: 2

5 Year Summary

Students graduated: 18

Scientific Publications: 75

Presentations: 93


Contact Us

Pennsylvania Cooperative Fish and Wildlife Research Unit University Park, PA 16802 Phone: (814) 865 - 4511 Fax: (814) 863 - 4710

Unit Leader

Duane Diefenbach
Duane R. Diefenbach

Upon completion of my Ph.D. I worked for the Pennsylvania Game Commission as their biometrician for 7 years. There I began my current research involving grassland sparrows and black bears. Since 1999 I have been a research scientist with the Pennsylvania Unit and enjoy the challenges in integrating research to improve management decisions.


  1. Penn State University Campus Maps

Pennsylvania Cooperative Fish and Wildlife Research Unit Cooperators

  1. Pennsylvania Fish and Boat Commission
  2. Pennsylvania Game Commission
  3. Pennsylvania State University
  4. U.S. Fish and Wildlife Service
  5. U.S. Geological Survey
  6. Wildlife Management Institute