Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Dr. Raymond Carthy

Research Publications

  • Rees, A. F., L. Avens, K. Ballorain, E. Bevan, A. C. Broderick, R. R. Carthy, M. J. Marjolijn, J. A. Christianen, G. Cuclos, M. R. Heithaus, D. W. Johnston, J . C. Mangel, F. Paladino, K. Pendoley, R. D. Reina, N. J. Robinson, R. Ryan, S. T. Sykora-Bodie, D. Tilley, M. R. Varela, E. R. Whitman, P. A. Whittock, T. Wibbels, and B. J. Godley. 2018.The potential of unmanned aerial systems for sea turtle research and conservation: a review and future directions. Endangered Species Research.
  • Bjorndal, K.A., Bolten, A.B., Chaloupka, M., Saba, V.S., Bellini, C., Marcovaldi, M.A.G., Santos, A.J.B., Bortolon, L.F.W., Meylan, A.B., Meylan, P.A., Gray, J., Hardy, R., Brost, B., Bresette, M., Gorham, J.C., Connett, S., Crouchley, B.V.S., Dawson, M., Hayes, D., Diez, C.E., van Dam, R.P., Willis, S., Nava, M., Hart, K.M., Cherkiss, M.S., Crowder, A.G., Pollock, C., Hillis-Starr, Z., Muñoz Tenería, F.A., Herrera-Pavón, R., Labrada-Martagón, V., Lorences, A., Negrete-Philippe, A., Lamont, M.M., Foley, A.M., Bailey, R., Carthy, R.R., Scarpino, R., McMichael, E., Provancha, J.A., Brooks, A., Jardim, A., López-Mendilaharsu, M., González-Paredes, D., Estrades, A., Fallabrino, A., Martínez-Souza, G., Vélez-Rubio, G.M., Boulon Jr., R.H., Collazo, J.A., Wershoven, R., Guzmán Hernández, V., Stringell, T.B., Sanghera, A., Richardson, P.B., Broderick, A.C., Phillips, Q., Calosso, M., Claydon, J.A.B., Metz, T.L., Gordon, A.L., Landry Jr., A.M., Shaver, D.J., Blumenthal, J., Collyer, L., Godley, B.J., McGowan, A., Witt, M.J., Campbell, C.L., Lagueux, C.J., Bethel, T.L., and Kenyon, L., 2017, Ecological regime shift drives declining growth rates of sea turtles throughout the West Atlantic. 2017. Global Change Biology, 23(11):4556-4568, Abstract |  Publisher Website | 
  • Phillips, A.C.N., J. Coutou, S. Rajh, N. Stewart, A. Watson, A. Jehu, H.Asmath, F. Dziva, C. Unakal R.B. Holder, and R.R. Carthy. 2017. Temporospatial dynamic and public health significance of bacterial flora identified on a major leatherback turtle (Dermochelys coriacea) nesting beach in the Southern Caribbean. Marine Ecology. 15pp IP- 068594.

    Abstract |  Publisher Website | 
  • Gallaway, Benny J. ,William J. Gazey, Charles W. Caillouet, Jr., Pamela T. Plotkin, Donna J. Shaver, F. Alberto Abreu Grobois, Anthony M. Amos, Patrick M. Burchfield, Ray R. Carthy, Marco A. Castro Martínez, John G. Cole, Andrew T. Coleman, Melissa Cook, Steven DiMarco, Sheryan P. Epperly, Allen M. Foley, Masami Fujiwara, Daniel Gomez Gamez, Gary L. Graham, Wade L. Griffin, Selina S. Heppell, Francisco Illescas Martínez, Jackie Isaacs, Margaret M. Lamont, Rebecca L. Lewison, Kenneth J. Lohmann, James M. Nance, Jonathan Pitchford, Nathan F. Putman, Scott W. Raborn, Jeffery K. Rester, Jack J. Rudloe, Laura Sarti Martínez, Mark Schexnayder, Jeffrey R. Schmid, Christopher Slay, Mandy Tumlin, Thane Wibbels and Blanca M. Zapata Najera. 2017. Development of a Kemp’s Ridley Sea Turtle Stock Assessment Model. Gulf of Mexico Science.

    Publisher Website | 
  • Carthy, R. R. 2016. The leatherback turtle: Biology and Conservation. Herpetological Review, 47(4):703-705.
  • Lamont, M.M., Fujisaki, I., Carthy, R.R. 2014. Estimates of vital rates for a declining loggerhead turtle (Caretta caretta) subpopulation: implications for management. Marine Biology. 161:2659–2668 DOI 10.1007/s00227-014-2537-0
  • Williams,N.C., Bjorndal, K. A., Lamont, M.M., Carthy, R.R. 2014. Winter diets of immature green turtles (Chelonia mydas) on a northern feeding ground: integrating stomach contents and stable isotope analyses. Estuaries and Coasts. 37:986-994. Download  | 
  • Carthy, R.R. The leatherbacks of Grande Riviere Trinidad and Tobago. 2012. Florida Environmental Outreach. (3), 1- 10-13. IP-034506
  • Nomani, S. Z., M. K. Oli, and R.R. Carthy. 2012. Line transects by design: the influence of study design, spatial distribution and density of objects on estimates of abundance. The Open Ecology Journal. 5: 25-44. Abstract | 
  • Shamblin, B.M., A.B. Bolten, K.A. Bjorndal, P.H. Dutton, J.T. Nielsen, F.A. Abreu-Grobois, K.J. Reich. B.E. Witherington, D.A. Bagley, L.M. Ehrhart, A.D. Tucker, D.S. Addison, A. Arenas, C. Johnson, R.R. Carthy, M.M. Lamont, M.G. Dodd, M.S. Gaines, E. LaCasella, and C.J. Nairn. 2012. Expanded mitochondrial control region sequences increase resolution of stock structure among North Atlantic loggerhead turtle rookeries. Marine Ecology Progress Series,. 469:145-160. Abstract | 
  • McMichael, E., R.R. Carthy, J.A. Seminoff and R.A. Scarpino. Kemp's ridley turtles in St. Joseph Bay, Florida, USA: Results from an in-water study in the northern Gulf of Mexico. NOAA Technical Memorandum NMFS-SEFSC-630
  • Salmon, M., R. Carthy, C. M.F. Lohmann, K. J. Lohmann, and J. Wyneken. 2012. Collecting a sample of loggerhead sea turtle hatchlings before a natural emergence does not reduce nest productivity. Endangered Species Research. 16: 296-299. doi: 10.3354/esr00409 IP-034556 Download  | 
  • Hart, K.M., M. Lamont, I. Fujisaki, A.D. Tucker, R.R. Carthy. Common coastal foraging areas in the Gulf of Mexico used by nesting loggerheads from 3 different subpopulations: Opportunities for marine conservation. Published.Biological Conservation. (145) 1:185-194. 2012.
  • Shamblin, B.M., M.G. Dodd, D.A. Bagley, L.M. Ehrhart, A.D. Tucker, C. Johnson, R.R. Carthy, R.A. Scarpino, E. McMichael, D.S. Addison, K.L. Williams, M.G. Frick, S. Ouellette, A.B. Meylan, M.H. Godfrey, S.R. Murphy, C.J. Nairn. Genetic structure of the southeastern United States loggerhead turtle nesting aggregation: evidence of additional structure within the peninsular Florida recovery unit. 2010. Marine Biology (2011) 158:571-587
  • Senko, J., V. Koch, W.M. Megill, R.R. Carthy, R.P. Templeton, W.J. Nichols. 2010. Fine scale daily movements and habitat use of East Pacific green turtles at a shallow coastal lagoon in Baja California Sur, Mexico. Journal of Experimental Marine Biology and Ecology. 391(2010): 92-100
  • Garland, K.A. and R.R. Carthy. 2010. Changing taste preferences, market demands and traditions in Pearl Lagoon, Nicaragua: a community reliant on Chelonia mydas for income and nutrition. Conservation and Society 8(1):55-72.
  • McMichael, E., J. Seminoff, R. Carthy. Kemp's ridley sea turtles in St. Joseph Bay, Florida, USA: results from an in-water study in the northeastern Gulf of Mexico. 2009. Journal of Natural History. 42 (43-44; 2293-2307).
  • Layton, J.E., T. Wibbels, T. Tucker, J. Wyneken, L. Ehrhart, R. Carthy, E.R. Martin, M. Bressette, C.Johnson, S. Fournier, and J. Schmid. 2009. Developing a Comprehensive Long-term Database on Nesting Beach Temperatures of the Loggerhead Sea Turtle in the Southeastern U.S: Applications and Implications for Global Climate Change. Integrative and Comparative Biology. 49:E227-E227 Publisher Website | 
  • McMichael, E., J. A. Seminoff and R. R. Carthy. Growth rates of wild green turtles, Chelonia mydas, at a temperate foraging habitat in the northern Gulf of Mexico: assessing short-term affects of cold stunning on growth. Journal of Natural History.
  • Nomani, S. Z., R. R. Carthy and M. K. Oli. Comparison of methods for estimating abundance of gopher tortoises. Applied Herpetology. 5:13-31
  • Lamont, M. M., and R. R. Carthy. 2007. Response of nesting sea turtles to barrier island dynamics. Chelonian Conservation and Biology 6(2): 206-212
  • Carthy, R. R. 2006. An owner's manual for the Chelonioidea. Conservation Biology. 20(1)259-260.
  • Gregory, C. J., R. R. Carthy, and L. G. Pearlstine. 2006. Application of Species at Risk at Camp Blanding Training Site, FLARNG(Florida). Southeastern Naturalist. 5(3):473-498.
  • Carthy, R. R., A. M. Foley, and Y. Matsuzawa. 2002. The incubation environment of nests of the loggerhead turtle (Caretta caretta) and effects of hatching success and the characteristics of hatchlings. Book Chapter, for "The Biology of the Loggerhead Sea Turtle". Smithsonian Institution Press, August 2003.
  • Tiwari, M., R. R. Carthy, A. Silveria, and K. A. Bjorndal. 2003. A comparison of nest architecture in loggerhead sea turtles (Caretta caretta) nesting in Florida and Brazil. Herpetological Review, Vol. 34(2):138-139.
  • Parris, L., M. M. Lamont, and R. R. Carthy. 2002. Increased incidence of red imported fire ant (Hymenoptera: formicidae) presence in loggerhead sea turtle (testudines: cheloniidae) nests and observations of hatchling mortality. Florida Entomologist, Vol. 85(3):514-517.
  • Askren, V., R. Carthy, C. Chase, G. Comstock, E. Franz and T. Oriola. 2002. More Citation Vigilance. The Scientist, Vol. 16(17):14.
  • Maglothin, M. L., M. M. Lamont, and R. R. Carthy. 2002. Nest Chamber anomalies in loggerhead sea turtle (Caretta caretta) nests from Northwest Florida, USA. Herpetological Review., Vol. 33(1):48-50.
  • Wooding, J. B., and R. R. Carthy. 2000. Interactions of humans and manatees in Crystal River, Florida. Proceedings of the Southeastern Association of Fish and Wildlife Agencies, 52:260-264.
  • Gross, T. S., D. A. Crain, K. A. Bjorndal, A. B. Bolten, and R. R. Carthy. 1995. Identification of sex hatchling loggerhead turtles (Caretta caretta) by analysis of steriod concentrations in chorioallantoic/amniotic fluid. Gen. Comp. Endocrinol. 99:204-210.
  • Carthy, R. R. 1992. Scanning electron microscopy (SEM) of loggerhead (Caretta caretta) eggshell structure. In: M. Salmon and J. Wyneken (compilers), Proceedings of the 11th Annual Workshop on Sea Turtle Biology and Conservation. NOAA Technical Memorandum NMFS-SEFSC-302:143-144.

Current Staff

Federal Staff: 2

Masters Students: 4

Phd Students: 5

Post Docs: 0

University Staff: 3

5 Year Summary

Students graduated: 8

Scientific Publications: 24

Presentations: 44


Contact Us

Florida Cooperative Fish and Wildlife Research Unit Gainesville, FL 32611-0485 Phone: (352) 846 - 0534 Fax: (352) 846 - 0841 Our University Web Site

Unit Leader

Abby Powell
Abby Powell

Abby joined the Florida Cooperative Fish and Wildlife Research Unit as Leader in 2015 and holds a joint appointment as Courtesy Professor in Wildlife Conservation and Ecology. She served as Assistant Unit Leader at the Alaska Unit 2000-2015.

Florida Cooperative Fish and Wildlife Research Unit Cooperators

  1. Florida Fish and Wildlife Conservation Commission
  2. U.S. Fish and Wildlife Service
  3. U.S. Geological Survey
  4. University of Florida
  5. Wildlife Management Institute