Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Peer Reviewed Publications

  • Ainley, DG, G Ballard, BJ Karl, and KM Dugger. 2005. Leopard seal predation rates at penguin colonies of different size. Antarctic Science 17:335-340. Publisher Website | 
  • Ainley, DG, G. Ballard, and KM Dugger. 2006. Competition among penguins and cetaceans reveals trophic cascades in the Western Ross Sea, Antarctica. Ecology 87:2080-2093.
  • Dugger, KM, G Ballard, DG Ainley, and K Barton. 2006. Effects of flipper-bands on apparent survival and foraging behavior of Adélie penguins. The Auk 123:858-869.
  • Ainley, DG, KM Dugger, V Toniolo, and I Gaffney. 2007. Cetacean occurrence patterns in the Amundsen and southern Bellingshausen Seas sector, Southern Ocean. Marine Mammal Science 23:287-305.
  • Lesroel, A, KM Dugger, G Ballard, and DG Ainley. 2009. Effects of individual quality, reproductive success and environmental variability on survival of a long-lived seabird. Journal of Animal Ecology 78:798-806.
  • Dugger, KM, DG Ainley, P Lyver, K B, and G Ballard. 2010. Survival differences and the effect of environmental instability on breeding dispersal in an Adélie penguin meta-population. Proceedings of the National Academy of Sciences 107:12375-12380.
  • Ballard, G, DG Ainley, KM Dugger, and N Nur. 2010. Foraging strategies of Adélie penguins: adjusting body condition to cope with environmental variability. Marine Ecology Progress Series 405:287-302.

  • LaRue, M.A., D.G. Ainley, M.Swanson, K.M. Dugger, P.O'B. Lyver, K.Barton, and G. Ballard. 2013. Climate change winners: receding ice fields facilitate colony expansion and altered dynamics in an Adélie penguin metapopulation. PLoS ONE 8(4):e60568. Doi:10.1371/journal.pone.0060568.
  • Dugger, K.M., G. Ballard, D.G. Ainley, P.O'B Lyver, and C. Schine. 2014. Adélie penguins coping with environmental change: results from a natural experiment at the edge of their breeding range. Frontiers in Ecology and Evolution.
  • Whitehead, A.L., P. O'B. Lyver, G. Ballard, K. Barton, B.J. Karl, K. M. Dugger, S. Jennings, A. Lescroel, P.R. Wilson, and D. G. Ainley. 2015. Factors driving Adelie penguin chick size, mass and condition at colonies of different sizes in the Southern Ross Sea. Marine Ecology Progress Series 523:199-213.
  • Jennings, S., A. Varsani, K.M. Dugger, G. Ballard, and D.G. Ainley. 2016. Sex-based differences in Adelie penguin (Pygoscelis adeliae) chick growth rates. PloS ONE 11(3): e0149090. doi:10.1371/
  • Wilson, D. J., P. O’B. Lyver, T. C. Greene, A. L. Whitehead, K. M. Dugger, B. J. Karl, J. R. F. Barringer, R. McGarry, A. M. Pollard, and D. G. Ainley. 2016. Adélie Penguin numbers predict breeding South Polar Skua numbers on Ross Island, Antarctica. Polar Biology early relesase: DOI 10.1007/s00300-016-1980-4
  • Youngflesh, C., S. Jenouvrier, Y.Li, R. Ji, D.G. Ainley, G. Ballard, C. Barbraud, K. Delord, K.M. Dugger, W.R. Fraser, J.T. Hinke, P.O’B. Lyver, S. Olmastroni, S.G. Trivelpiece, W.Z. Trivelpiece, and H.J. Lynch. Importance of environmental variability in phenological match-mismatch: lessons from a circumpolar analysis of the Adélie penguin. Ecology 98(4):940-951.
  • Van Doorslaer, K, V. Ruoppolo, A. Schmidt, A. Lescroel, D. Jongsomjit, M. Elrod, S. Kraberger, D. Stainton, K.M. Dugger, G. Ballard, D.G. Ainley, and A. Varsani. 2017.
    Unique genome organization of non-mammalian
    papillomaviruses provides insights into the evolution
    of viral early proteins. Viral Evolution 3(2):1-12. doi: 10.1093/ve/vex027
  • Gremillet, D., A. Lescroel, G. Ballard, K.M. Dugger, M. Massaro, and D.G. Ainley. 2018. Energetic fitness: Field metabolic rates assessed via 3D
    accelerometry complement conventional fitness metrics. Functional Ecology 32:1203-1213.
  • Ainley, D.G., K.M. Dugger, M. La Mesa, G. Ballard, K.J. Barton, S. Jennings, B.J. Karl, A. Lescroël, P.O’B. Lyver, A. Schmidt, P. Wilson. 2018. Post-fledging survival of Adélie penguins at multiple colonies: Chicks raised
    on fish do well. Marine Ecology Progress Series 601:239-251. DOI: 10.3354/meps12687

Current Staff

Federal Staff: 98

Masters Students: 230

Phd Students: 144

Post Docs: 54

University Staff: 248

5 Year Summary

Students graduated: 647

Scientific Publications: 1894

Presentations: 4159


Featured Peer Publication

Silver carp are skittish around boats - especially when the boat is pumping out electrical current Jumping Silver Carp


  1. AFS Journals
  2. TWS Publications

Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey