Cooperative Research Units
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Export of Invertebrate Drift From Headwater Streams

Headwaters stream


January 2015 - June 2018


This study is part of a larger multidisciplinary project investigating stream ecosystem response to riparian management an includes scientists from Green Diamond Resource Company, Humboldt State University, Oregon State University and the U.S. Forest Service. The study will be conducted on Green Diamond lands in tributaries of the lower Klamath River in coastal northern California. Sites will be chosen that differ in percentage of coniferous riparian trees, but which are similar in confluence with a fish-bearing reach.
Objectives of this study are to: 1) Quantify the magnitude, composition, and energy content of invertebrate subsidies of terrestrial and aquatic origin from Class II (fish-less) streams to downstream fish-bearing waters at different times of year and from streams differing in riparian canopy; and 2) Evaluate use of these invertebrate subsidies by fish and their potential contribution to fish growth.


Current Staff

Federal Staff: 95

Masters Students: 239

Phd Students: 142

Post Docs: 53

University Staff: 240

5 Year Summary

Students graduated: 701

Scientific Publications: 1936

Presentations: 4260



  • Margaret WilzbachPrincipal Investigator
  • Jon HollisStudent

Funding Agencies

  • Green Diamon Resources Company


Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey