Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Development and testing of a protocol for monitoring mussels


August 2005 - December 2009


1. Evaluate alternative sampling designs for determining the species composition, density, and recruitment of the
mussel assemblage with the Mammoth Cave National Park
2. Develop a sampling protocol for the continued long-term monitoring of the assemblage
3. Evaluate the effectiveness and efficiency of the sampling protocol


Current Staff

Federal Staff: 99

Masters Students: 225

Phd Students: 143

Post Docs: 53

University Staff: 247

5 Year Summary

Students graduated: 658

Scientific Publications: 1872

Presentations: 4186



  • James LayzerPrincipal Investigator

Funding Agencies

  • Leetown Science Center

Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey