Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Development and testing of a protocol for monitoring mussels


August 2005 - December 2009


1. Evaluate alternative sampling designs for determining the species composition, density, and recruitment of the
mussel assemblage with the Mammoth Cave National Park
2. Develop a sampling protocol for the continued long-term monitoring of the assemblage
3. Evaluate the effectiveness and efficiency of the sampling protocol


Current Staff

Federal Staff: 95

Masters Students: 244

Phd Students: 144

Post Docs: 55

University Staff: 251

5 Year Summary

Students graduated: 680

Scientific Publications: 1916

Presentations: 4252



  • James LayzerPrincipal Investigator

Funding Agencies

  • Leetown Science Center

Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey