Cooperative Research Units
Education, Research And Technical Assistance For Managing Our Natural Resources
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Gopher Tortoise (gopherus polyphermus)Population Survey For St. Marks NWR-Line Transect Distance Sampling


September 2010 - August 2011


The gopher tortoise (Gopherus polyphemus), unlisted range, is among the U.S. Fish and Wildlife Ecological Services 2008 priorities as a “preclude the need to list” species. This project furthers the Strategic Habitat Conservation process by providing critical information necessary for outcome-based monitoring, planning and management at St. Marks National Wildlife Refuge.


Current Staff

Federal Staff: 95

Masters Students: 239

Phd Students: 142

Post Docs: 53

University Staff: 240

5 Year Summary

Students graduated: 701

Scientific Publications: 1936

Presentations: 4260



  • Raymond CarthyPrincipal Investigator
  • Margaret LamontCo-Principal Investigator

Funding Agencies

  • US Geological Survey


Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey