Cooperative Research Units
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Life history of the Cumberland papershell (anodontoides denigrata)


September 2011 - December 2014


The Cumberland papershell (Anodontoides denigrata) is a little known species. It was originally described by Lea in 1852, but was later considered a synonym of A. ferussacianus and forgotten about until Cicerello and Schuester recognized it as a distinct species in 2003. It is known from only a few small tributaries of the Big South Fork in TN and the upper Cumberland River, KY. Virtually nothing is known about the life history of the Cumberland papershell. This study will focus on describing its life history traits. Specific objectives will include determining: at what age and size individuals become reproductively mature, when spawning occurs and how long larvae are brooded, fecundity, and what host fishes are needed to complete its life cycle.


Current Staff

Federal Staff: 95

Masters Students: 238

Phd Students: 144

Post Docs: 54

University Staff: 239

5 Year Summary

Students graduated: 673

Scientific Publications: 1905

Presentations: 4235



  • James LayzerPrincipal Investigator
  • Tyler HernStaff

Funding Agencies

  • Tennessee Field office


Cooperative Research Units Program Headquarters Cooperators

  1. U.S. Geological Survey